Just as Gio makes Betty an incredibly romantic offer, Henry pops back into her life with a proposal of his own. Meanwhile, Betty finds herself centerfield at the charity softball game of the year: Mode vs. ELLE Magazine. Can Wilhelmina strike out ELLE’s star player, supermodel Naomi Campbell? In other developments, an orphaned boy from France turns up at Mode with an amazing claim, and Hilda learns some shocking news about Coach Diaz.
Betty, still missing Henry, fights her attraction to Gio, but later finds herself stuck with him when both volunteer to chaperone Justin’s middle school dance. Meanwhile, Hilda tries to get Coach Diaz (guest star Eddie Cibrian) to notice her; Amanda agrees to do a reality show with her dad, KISS rock legend Gene Simmons; and Daniel feels the heat when Wilhelmina makes her triumphant return to Mode amid a media frenzy.
Betty switches gears with rival Charlie (guest star Jayma Mays) and tries to befriend her by throwing a baby shower, a party that no one will ever forget. Meanwhile, Hilda confronts Justin’s PE teacher (guest star Eddie Cibrian as Coach Diaz) when she learns Justin is flunking gym; Daniel unravels after his break-up with Renee, and Christina finally learns who fathered the baby she’s carrying for Wilhelmina.
An increasingly delusional Renee (guest star Gabrielle Union) becomes convinced that Betty is in love with Daniel and therefore is her rival for his affections. Meanwhile, Hilda’s new client at the beauty shop turns out to be ex-neighbor and lifetime nemesis Gina Gambarro (guest star Ava Gaudet), who returns to Queens to rub her fabulous life and wealthy doctor husband in Hilda’s face.
Betty’s plans to spend a romantic 24th birthday with Henry are sidelined when his Baby Mama Charlie drops in unexpectedly. Will Gio be able to salvage Betty’s big day? Meanwhile, Wilhelmina does her devious best to spoil Renee’s blossoming relationship with Daniel by hinting there’s some dark secrets in her sister’s past.
Betty gets an assignment from Daniel to interview “Phil Roth,” thinking it’s a sit down with the great novelist, but later learning this Phil Roth is a writer who has penned a series of books about picking up women. Meanwhile Daniel hooks up with a sexy new lady, Renee, only to learn they have a shocking connection to each other, and Amanda and Marc hatch a plan to reach out to the man they believe is Amanda’s biological dad.
Betty’s new perfume is driving Henry wild and Betty crazy, literally, after she unwittingly begins using a deadly scent concocted by the late Fey Sommers and starts acting very strangely. Meanwhile, Meade matriarch Claire goes on the stand for Fey’s murder, a vindictive Amanda withholds important information, and Wilhelmina takes steps to become a member of the Meade clan — one way or the other.
Amanda visits a psychic (guest star Annie Potts) for insight into her dad’s identity. Meanwhile, disturbed about the message Mode is sending to Justin and his classmates after they visit the magazine, Betty pushes Daniel to use “real” models during fashion week; and Wilhelmina gets some unexpected news about her latest plans for the Meade dynasty.
Henry reluctantly agrees to go on a double date with Betty and an unlikely couple — Hilda and Gio. Meanwhile, Wilhelmina’s attempt to change her image as the queen of mean fails miserably after she tangles with America’s most beloved golden girl.
A Wilhelmina-induced crisis at Mode finds everyone scrambling to re-create an issue on deadline. Desperate times call for desperate measures, as Daniel and Betty try to break wild child cover girl (Eliza Dushku as Cameron Ashlock) out of rehab for a re-shoot, and Alexis tries to charm the printer (guest star David J. Steinberg) into extending their time. Meanwhile, Betty struggles with an unwelcome (and persistent) visitor who hopes to goad her into making good on a promise.